Haeterini

Niklas Wahlberg

---

Updated 2018-01-16. Based on

Lamas, G. 2004: Atlas of Neotropical Lepidoptera. Checklist: Part 4A, Hesperioidea - Papilionoidea. Scientific Publishers, Gainsville, Florida.

Penz, C. M., Alexander, L. G. & DeVries, P. J. 2014: Revised species definitions and nomenclature of the rose colored Cithaerias butterflies (Lepidoptera, Nymphalidae, Satyrinae). Zootaxa 3873: 541-559.

The Butterflies of America

TRIB Haeterini

GENU Cithaerias Hübner, [1819]

GSYN Callitaera Butler, 1868

SPEC Cithaerias andromeda (Fabricius, 1775)

SUBS Cithaerias andromeda andromeda (Fabricius, 1775)

ORIG Papilio andromeda Fabricius, 1775
SSYN Papilio philis Stoll, 1782
SSYN Cithaerias cissa Hübner, [1819]
SSYN Hetaera [sic] pellucida Butler, 1866
SSYN Hetaera [sic] phelis Butler, 1866
SSYN Hetaera [sic] harpalyce Butler, 1866
SSYN Callitaera pellucida f. archeops Le Cerf, 1927
SSYN Callitaera philis f. hyacinthina Langer, 1943

SUBS Cithaerias andromeda azurina (J. Zikán, 1942)

ORIG Callitaera andromeda azurina J. Zikán, 1942

SUBS Cithaerias andromeda bandusia Staudinger, [1887]
SUBS Cithaerias andromeda esmeralda (Doubleday, 1845)

ORIG Haetera esmeralda Doubleday, 1845
SSYN Callitaera rubina Fassl, 1922

SPEC Cithaerias cliftoni Constantino, 1995
SPEC Cithaerias aurora (C. Felder & R. Felder, 1862)

SUBS Cithaerias aurora aurora (C. Felder & R. Felder, 1862)

ORIG Haetera aurora C. Felder & R. Felder, 1862
SSYN Callitaera phantoma Fassl, 1922
SSYN Callitaera aura Langer, 1943
SSYN Cithaerias juruaënsis [sic] d'Almeida, 1951

SUBS Cithaerias aurora tambopata Lamas, 1998

SPEC Cithaerias aurorina (Weymer, 1910)

ORIG Callitaera aurorina Weymer, 1910
SSYN Callitaera merolina J. Zikán, 1942

SPEC Cithaerias pireta (Stoll, 1780)

SUBS Cithaerias pireta pireta (Stoll, 1780)

ORIG Papilio pireta Stoll, 1780
SSYN Papilio menander Drury, 1782

SUBS Cithaerias pireta magdalenensis Constantino, 1995

SPEC Cithaerias pyritosa (J. Zikán, 1942)

ORIG Callitaera pyritosa J. Zikán, 1942
SSYN Cithaerias similigena d'Almeida, 1951
SSYN Callitaera pireta amaryllis (Bryk, 1953)

SPEC Cithaerias pyropina (Salvin & Godman, 1868)

SUBS Cithaerias pyropina pyropina (Salvin & Godman, 1868)

ORIG Callitaera pyropina Salvin & Godman, 1868
SSYN Callitaera pyropina var. rosacea Langer, 1944
SSYN Callitaera pyropina roquensis Bryk, 1953

SUBS Cithaerias pyropina songoana (Langer, 1944)

ORIG Callitaera songoana Langer, 1944

SUBS Cithaerias pyropina julia Willmott, 2015

GENU Dulcedo d'Almeida, 1951

SPEC Dulcedo polita (Hewitson, 1869)

ORIG Haetera polita Hewitson, 1869

GENU Haetera Fabricius, 1807

GSYN Oreas Hübner, [1807]
GSYN Maniola Rafinesque, 1815
GSYN Hetaera Hoffmannsegg, 1818 (unj. emend. Haetera Fabricius, 1807)
GSYN Pselna Billberg, 1820 (unn. repl. Haetera Fabricius, 1807)

SPEC Haetera macleannania H.W. Bates, 1865

SSYN Haetera macleannania laddeya F.M. Brown, [1943]
SSYN Haetera macleannania laddeya f. neglecta F.M. Brown, [1943]

SPEC Haetera piera (Linnaeus, 1758)

SUBS Haetera piera piera (Linnaeus, 1758)

ORIG Papilio piera Linnaeus, 1758

SUBS Haetera piera diaphana Lucas, 1857

SSYN Haetera hymenaea C. Felder & R. Felder, 1867

SUBS Haetera piera negra C. Felder & R. Felder, 1862

SSYN Haetera piera f. ecuadora F.M. Brown, [1943]
SSYN Haetera piera f. pseudopiera F.M. Brown, [1943]
NUDU Haetera pirea [sic] f. mariuá [sic] W. Zikán & Wygodzinsky, 1948
SSYN Haetera piera lesbia Bryk, 1953
SSYN Haetera piera lesbia f. metathetica Bryk, 1953

SUBS Haetera piera pakitza Lamas, 1998
SUBS Haetera piera unocellata Weymer, 1910

GENU Pierella Westwood, 1851

GSYN Pieris Hübner, [1819]
GSYN Antirrhaea Westwood, 1851

SPEC Pierella amalia Weymer, 1885

SSYN Pierella lena ab. leucospila Staudinger, 1887

SPEC Pierella astyoche (Erichson, [1849])

SUBS Pierella astyoche astyoche (Erichson, [1849])

ORIG Hetaera [sic] astyoche Erichson, [1849]
SSYN Haetera larymna Doubleday, [1849]

SUBS Pierella astyoche bernhardina Bryk, 1953

NUDU Pierella astyoche f. obscura W. Zikán & Wygodzinsky, 1948

SUBS Pierella astyoche stollei Ribeiro, 1931

SPEC Pierella helvina (Hewitson, 1859)

SUBS Pierella helvina helvina (Hewitson, 1859)

ORIG Haetera helvina Hewitson, 1859

SUBS Pierella helvina hymettia Staudinger, [1886]

SSYN Pierella incanescens werneri Hering & Hopp, 1925

SUBS Pierella helvina incanescens Godman & Salvin, 1877

SSYN Pierella incanescens costaricana Niepelt, 1927

SUBS Pierella helvina ocreata Salvin & Godman, 1868

NUDU Pierella ocreata Butler, 1868,

SUBS Pierella helvina pacifica Niepelt, 1924

SSYN Pierello [sic] ocreata johnsoni Talbot, 1932

SPEC Pierella hortona (Hewitson, 1854)

SUBS Pierella hortona hortona (Hewitson, 1854)

ORIG Haetera hortona Hewitson, 1854
SSYN Haetera hortensia C. Felder & R. Felder, 1862
SSYN Pierella hortona f. albopunctata F.M. Brown, 1948
SSYN Pierella hortona f. ocellata Smart, 1975

SUBS Pierella hortona albofasciata Rosenberg & Talbot, 1914

SSYN Pierella albofaciata [sic] decepta F.M. Brown, 1948

SPEC Pierella hyalinus (Gmelin, [1790])

SUBS Pierella hyalinus hyalinus (Gmelin, [1790])

ORIG Papilio hyalinus Gmelin, [1790]
SSYN Pieris dracontis Hübner, [1819]
SSYN Pierella hyalinus fusimaculata F.M. Brown, 1948

SUBS Pierella hyalinus extincta Weymer, 1910
SUBS Pierella hyalinus schmidti Constantino, 1995
SUBS Pierella hyalinus velezi Constantino, 1995

SPEC Pierella hyceta (Hewitson, 1859)

SUBS Pierella hyceta hyceta (Hewitson, 1859)

ORIG Haetera hyceta Hewitson, 1859

SUBS Pierella hyceta ceryce (Hewitson, 1874)

ORIG Haetera ceryce Hewitson, 1874

SUBS Pierella hyceta latona (C. Felder & R. Felder, 1867)

ORIG Haetera latona C. Felder & R. Felder, 1867

SPEC Pierella lamia (Sulzer, 1776)

SUBS Pierella lamia lamia (Sulzer, 1776)

ORIG Papilio lamia Sulzer, 1776
OSYN Papilio rhea Fabricius, 1775
SSYN Papilio dyndimene Cramer, 1779
SSYN Pierella luna ab. albina Oberthür, 1896
SSYN Pierella lamia f. fabriciana Ebert, 1965

SUBS Pierella lamia boliviana F.M. Brown, 1948
SUBS Pierella lamia chalybaea Godman, 1905

SSYN Haetera lamia f. columbina E. Krüger, 1925

SPEC Pierella lena (Linnaeus, 1767)

SUBS Pierella lena lena (Linnaeus, 1767)

ORIG Papilio lena Linnaeus, 1767
SSYN Papilio sectator (Meerburgh, 1775)

SUBS Pierella lena brasiliensis (C. Felder & R. Felder, 1862)

ORIG Haetera brasiliensis C. Felder & R. Felder, 1862
SSYN Pierella lena glaucolena Weymer, 1910
NUDU Pierella lena f. melanosa W. Zikán & Wygodzinsky, 1948
SSYN Pierella lena f. obsoleta F.M. Brown, 1948
SSYN Pierella lena browni Forster, 1964

SUBS Pierella lena salma Constantino, 2007

SPEC Pierella lucia Weymer, 1885

SSYN Pierella astyoche var. albomaculata Staudinger, 1887

SPEC Pierella luna (Fabricius, 1793)

SUBS Pierella luna luna (Fabricius, 1793)

ORIG Papilio luna Fabricius, 1793
SSYN Pierella luna f. rubra F.M. Brown, 1948

SUBS Pierella luna lesbia Staudinger, 1887
SUBS Pierella luna pallida (Salvin & Godman, 1868)

ORIG Hetaera [sic] pallida Salvin & Godman, 1868

SUBS Pierella luna rubecula Salvin & Godman, 1868

SSYN Haetera heracles Boisduval, 1870

SPEC Pierella nereis (Drury, 1782)

ORIG Papilio nereis Drury, 1782

GENU Pseudohaetera Brown, [1943]

GSYN Paradulcedo Constantino, 1992

SPEC Pseudohaetera hypaesia (Hewitson, 1854)

ORIG Haetera hypaesia Hewitson, 1854
NUDU Haetera hippomene Boisduval, 1870
SSYN Haetera hippomene f. strandi (Niepelt, 1922)

SPEC Pseudohaetera mimica (Rosenberg & Talbot, 1914)

ORIG Callitaera mimica Rosenberg & Talbot, 1914
SSYN Haetera hypaesia var. obscura E. Krüger, 1925
NUDU Hetaera [sic] signata E. Krüger, 1933
SSYN Cithaerias gilmouri Okano, 1986


---

Back to The Classification of Nymphalidae