Heliconiini

Niklas Wahlberg

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Updated 2022-08-01. Based on

• The Butterflies of America
• Farfán J, Cerdeña J, Vargas HA, Gonçalves GL, Lamas G, Moreira GRP 2022: A peculiar new species of Dione (Agraulis) Boisduval & Le Conte (Lepidoptera, Nymphalidae, Heliconiinae) associated with Malesherbia Ruiz & Pavón (Passifloraceae) in xeric western slopes of the Andes. ZooKeys 1113: 199-226. doi:10.3897/zookeys.1113.85769
• Kozak, K. M., Wahlberg, N., Neild, A., Dasmahapatra, K., Mallet, J. & Jiggins, C. D. 2015: Multilocus species trees show the recent adaptive radiation of the mimetic Heliconius butterflies. Systematic Biology 64: 505-524. doi:10.1093/sysbio/syv007
• Nunez et al. 2021: Integrative taxonomy clarifies species limits in the hitherto monotypic passion-vine butterfly genera Agraulis and Dryas (Lepidoptera, Nymphalidae, Heliconiinae). Systematic Entomology doi:10.1111/syen.12523
• Rosser, N., Freitas, A. V. L., Huertas, B., Joron, M., Lamas, G., Mérot, C., Simpson, F., Willmott, K. R., Mallet, J. & Dasmahaptra, K. K. 2018: Cryptic speciation associated with geographic and ecological divergence in two Amazonian Heliconius butterflies. Zoological Journal of the Linnean Society doi:10.1093/zoolinnean/zly046

TRIB Heliconiini Swainson, 1822

GENU Agraulis Boisduval & Le Conte, [1835]

SPEC Agraulis vanillae (Linnaeus, 1758)
SPEC Agraulis incarnata (N. Riley, 1926)

SUBS Agraulis incarnata incarnata (N. Riley, 1926)
SUBS Agraulis incarnata nigrior Michener, 1942

SPEC Agraulis forbesi Michener, 1942
SPEC Agraulis insularis Maynard, 1889
SPEC Agraulis maculosa (Stichel, [1908])
SPEC Agraulis lucina C. Felder & R. Felder, 1862
SPEC Agraulis galapagensis W. Holland, 1890
SPEC Agraulis dodona Lamas & Farfán, 2022

GENU Dione Hübner, [1819]

SPEC Dione moneta Hübner, [1825]

SUBS Dione moneta poeyii Butler, 1873
SUBS Dione moneta butleri Stichel, [1908]
SUBS Dione moneta moneta Hübner, [1825]

SPEC Dione glycera (C. Felder & R. Felder, 1861)
SPEC Dione juno (Cramer, 1779)

SUBS Dione juno huascuma (Reakirt, 1866)
SUBS Dione juno juno (Cramer, 1779)
SUBS Dione juno andicola (H. Bates, 1864)
SUBS Dione juno miraculosa M. Hering, 1926
SUBS Dione juno suffumata K. Brown & O. Mielke, 1972

GENU Podotricha Michener, 1942

SPEC Podotricha judith (Guérin-Méneville, [1844])

SUBS Podotricha judith judith (Guérin-Méneville, [1844])
SUBS Podotricha judith caucana (N. Riley, 1926)
SUBS Podotricha judith mellosa (Stichel, 1906)
SUBS Podotricha judith straminea (N. Riley, 1926)

SPEC Podotricha telesiphe (Hewitson, 1867)

SUBS Podotricha telesiphe telesiphe (Hewitson, 1867)
SUBS Podotricha telesiphe tithraustes (Salvin, 1871)

GENU Dryadula Michener, 1942

SPEC Dryadula phaetusa (Linnaeus, 1758)

GENU Dryas Hübner, [1807]

GSYN Colaenis Hübner, 1819

SPEC Dryas iulia (Fabricius, 1775)

SUBS Dryas iulia iulia (Fabricius, 1775)
SUBS Dryas iulia largo Clench, 1975
SUBS Dryas iulia nudeola (M. Bates, 1934)
SUBS Dryas iulia carteri (N. Riley, 1926)
SUBS Dryas iulia zoe L. Miller & Steinhauser, 1992
SUBS Dryas iulia delila (Fabricius, 1775)
SUBS Dryas iulia fucatus (Boddaert, 1783)

SPEC Dryas dominicana (A. Hall, 1917)

SUBS Dryas dominicana dominicana (A. Hall, 1917)
SUBS Dryas dominicana warneri (A. Hall, 1936)

SPEC Dryas lucia (N. Riley, 1926)

SUBS Dryas lucia lucia (N. Riley, 1926)
SUBS Dryas lucia martinica Enrico & Pinchon, 1969

SPEC Dryas alcionea (Cramer, 1779)

SUBS Dryas alcionea alcionea (Cramer, 1779)
SUBS Dryas alcionea moderata (N. Riley, 1926)
SUBS Dryas alcionea framptoni (N. Riley, 1926)

GENU Philaethria Billberg, 1820

SPEC Philaethria diatonica (Fruhstorfer, 1912)
SPEC Philaethria dido (Linnaeus, 1763)

SUBS Philaethria dido panamensis Constantino & Salazar, 2010
SUBS Philaethria dido chocoensis Constantino, 1999
SUBS Philaethria dido dido (Linnaeus, 1763)

SPEC Philaethria constantinoi Salazar, 1991
SPEC Philaethria browni Constantino & Salazar, 2010
SPEC Philaethria romeroi Constantino & Salazar, 2010
SPEC Philaethria neildi Constantino & Salazar, 2010

SUBS Philaethria neildi winhardi Constantino & Salazar, 2010
SUBS Philaethria neildi neildi Constantino & Salazar, 2010
SUBS Philaethria neildi tachiraensis Constantino & Salazar, 2010

SPEC Philaethria ostara (Röber, 1906)

SUBS Philaethria ostara araguensis Constantino & Salazar, 2010
SUBS Philaethria ostara meridensis Constantino & Salazar, 2010
SUBS Philaethria ostara ostara (Röber, 1906)

SPEC Philaethria andrei Brévignon, 2002

SUBS Philaethria andrei orinocensis Constantino & Salazar, 2010
SUBS Philaethria andrei andrei Brévignon, 2002

SPEC Philaethria pygmalion (Fruhstorfer, 1912)

SUBS Philaethria pygmalion metaensis Constantino & Salazar, 2010
SUBS Philaethria pygmalion pygmalion (Fruhstorfer, 1912)

SPEC Philaethria wernickei (Röber, 1906)

GENU Eueides Hübner, 1816

GSYN Mechanitis Illiger, 1807
GSYN Semelia Doubleday, 1844
GSYN Evides Agassiz, 1846
GSYN Semelia Erichson, 1848 (preocc. Semelia Doubleday, 1844)
GSYN Semelia Boisduval, 1870 (preocc. Semelia Doubleday, 1844)
GSYN Eisides Scudder, 1875

SPEC Eueides aliphera (Godart, 1819)

SUBS Eueides aliphera aliphera (Godart, 1819)
SUBS Eueides aliphera cyllenella Seitz, 1912
SUBS Eueides aliphera gracilis Stichel, 1903

SPEC Eueides emsleyi K. Brown, 1976

SUBS Eueides emsleyi emsleyi K. Brown, 1976
SUBS Eueides emsleyi esmeraldensis Cast, 2013

SPEC Eueides heliconioides C. Felder & R. Felder, 1861

SUBS Eueides heliconioides heliconioides C. Felder & R. Felder, 1861
SUBS Eueides heliconioides eanes Hewitson, 1861
SUBS Eueides heliconioides koenigi H. Holzinger & R. Holzinger, 1993

SPEC Eueides isabella (Stoll, 1781)

SUBS Eueides isabella isabella (Stoll, 1781)
SUBS Eueides isabella arquata Stichel, 1903
SUBS Eueides isabella cleobaea Geyer, 1832
SUBS Eueides isabella dianasa (Hübner, [1806])
SUBS Eueides isabella dissoluta Stichel, 1903
SUBS Eueides isabella dynastes C. Felder & R. Felder, 1861
SUBS Eueides isabella ecuadorensis Strand, 1912
SUBS Eueides isabella eva (Fabricius, 1793)
SUBS Eueides isabella hippolinus Butler, 1873
SUBS Eueides isabella huebneri Ménétriés, 1857
SUBS Eueides isabella melphis (Godart, 1819)
SUBS Eueides isabella nigricornis R. G. Maza, 1982

SPEC Eueides lampeto Bates, 1862

SUBS Eueides lampeto lampeto Bates, 1862
SUBS Eueides lampeto acacetes Hewitson, 1869
SUBS Eueides lampeto apicalis Röber, 1927
SUBS Eueides lampeto brownsbergensis Gernaat & Beckles, 2010
SUBS Eueides lampeto carbo Seitz, 1913
SUBS Eueides lampeto concisa Lamas, 1985
SUBS Eueides lampeto nigrofulva Kaye, 1906

SPEC Eueides libitina Staudinger, 1885

SUBS Eueides libitina libitina Staudinger, 1885
SUBS Eueides libitina malleti Lamas, 1998

SPEC Eueides lineata Salvin & Godman, 1868
SPEC Eueides lybia (Fabricius, 1775)

SUBS Eueides lybia lybia (Fabricius, 1775)
SUBS Eueides lybia lybioides Staudinger, 1876
SUBS Eueides lybia olympia (Fabricius, 1793)
SUBS Eueides lybia orinocensis K. Brown & F. Fernández, 1985
SUBS Eueides lybia otelloi K. Brown & F. Fernández, 1985
SUBS Eueides lybia salcedoi K. Brown & F. Fernández, 1985

SPEC Eueides pavana Ménétriés, 1857
SPEC Eueides procula E. Doubleday, [1847]

SUBS Eueides procula procula E. Doubleday, [1847]
SUBS Eueides procula asidia Schaus, 1920
SUBS Eueides procula browni H. Holzinger & R. Holzinger, 1974
SUBS Eueides procula edias Hewitson, 1861
SUBS Eueides procula eurysaces Hewitson, 1864
SUBS Eueides procula kuenowii Dewitz, 1877
SUBS Eueides procula vulgiformis Butler & H. Druce, 1872

SPEC Eueides tales (Cramer, 1775)

SUBS Eueides tales tales (Cramer, 1775)
SUBS Eueides tales barcellinus J. Zikán, 1937
SUBS Eueides tales calathus Stichel, 1909
SUBS Eueides tales cognata Weymer, 1890
SUBS Eueides tales franciscus K. Brown & H. Holzinger, 1973
SUBS Eueides tales michaeli J. Zikán, 1937
SUBS Eueides tales pseudeanes Boullet & Le Cerf, 1910
SUBS Eueides tales pythagoras W. F. Kirby, [1900]
SUBS Eueides tales surdus Stichel, 1903
SUBS Eueides tales tabernula Lamas, 1985
SUBS Eueides tales xenophanes C. Felder & R. Felder, 1865

SPEC Eueides vibilia (Godart, 1819)

SUBS Eueides vibilia vibilia (Godart, 1819)
SUBS Eueides vibilia mereaui (Hübner, 1823)
SUBS Eueides vibilia unifasciatus Butler, 1873
SUBS Eueides vibilia vialis Stichel, 1903
SUBS Eueides vibilia vicinalis Stichel, 1903

GENU Heliconius Kluk, 1780

GSYN Migonitis Hübner, 1816
GSYN Sunias Hübner, 1816
GSYN Sicyonia Hübner, 1816
GSYN Ajantis Hübner, 1816
GSYN Apostraphia Hübner, 1816
GSYN Crenis Hübner, 1821
GSYN Phlogris Hübner, 1825
GSYN Blanchardia Buchecker, 1880 (preocc. Blanchardia Castelnau, 1875)
GSYN Laparus Billberg, 1820
GSYN Neruda Turner, 1976

SPEC Heliconius aoede (Hübner, [1813])

SUBS Heliconius aoede aoede (Hübner, [1813])
SUBS Heliconius aoede aliciae Neukirchen, 2000
SUBS Heliconius aoede astydamia (Erichson, [1849])
SUBS Heliconius aoede auca Neukirchen, 1997
SUBS Heliconius aoede ayacuchensis (Neukirchen, 1992)
SUBS Heliconius aoede bartletti (H. Druce, 1876)
SUBS Heliconius aoede centurius (Neukirchen, 1994)
SUBS Heliconius aoede cupidineus (Stichel, 1906)
SUBS Heliconius aoede emmelina (Oberthür, 1902)
SUBS Heliconius aoede eurycleia (K. Brown, 1973)
SUBS Heliconius aoede faleria (Fruhstorfer, 1910)
SUBS Heliconius aoede lucretius (Weymer, 1891)
SUBS Heliconius aoede manu Lamas, 1976
SUBS Heliconius aoede philipi (K. Brown, 1976)

SPEC Heliconius metis (Moreira & Mielke, 2010)
SPEC Heliconius godmani (Staudinger, 1882)
SPEC Heliconius metharme (Erichson, [1849])

SUBS Heliconius metharme metharme (Erichson, [1849])
SUBS Heliconius metharme makiritare K. Brown & F. Fernández, 1985
SUBS Heliconius metharme perseis (Stichel, 1923)

SPEC Heliconius antiochus (Linnaeus, 1767)

SUBS Heliconius antiochus antiochus (Linnaeus, 1767)
SUBS Heliconius antiochus aranea (Fabricius, 1793)
SUBS Heliconius antiochus araneides Staudinger, 1897
SUBS Heliconius antiochus salvinii Dewitz, 1877

SPEC Heliconius astraea Staudinger, 1897

SUBS Heliconius astraea astraea Staudinger, 1897
SUBS Heliconius astraea rondonia K. Brown, 1973

SPEC Heliconius atthis (E. Doubleday, 1847)
SPEC Heliconius besckei (Ménétriés, 1857)
SPEC Heliconius burneyi (Hübner, [1831])

SUBS Heliconius burneyi burneyi (Hübner, [1831])
SUBS Heliconius burneyi ada Neustetter, 1925
SUBS Heliconius burneyi anjae Neukirchen, 1995
SUBS Heliconius burneyi boliviensis Neukirchen, 1995
SUBS Heliconius burneyi catharinae Staudinger, 1885
SUBS Heliconius burneyi huebneri Staudinger, 1897
SUBS Heliconius burneyi jamesi Neukirchen, 1995
SUBS Heliconius burneyi koenigi Neukirchen, 1995
SUBS Heliconius burneyi lindigii C. Felder & R. Felder, 1865
SUBS Heliconius burneyi skinneri K. Brown & F. Fernández, 1985

SPEC Heliconius charithonia (Linnaeus, 1767)

SUBS Heliconius charithonia vazquezae W. Comstock & F. Brown, 1950
SUBS Heliconius charithonia tuckeri W. Comstock & F. Brown, 1950
SUBS Heliconius charithonia ramsdeni W. Comstock & F. Brown, 1950
SUBS Heliconius charithonia simulator Röber, 1921
SUBS Heliconius charithonia churchi W. Comstock & F. Brown, 1950
SUBS Heliconius charithonia charithonia (Linnaeus, 1767)
SUBS Heliconius charithonia antiquus Lamas, 1988
SUBS Heliconius charithonia bassleri W. Comstock & F. Brown, 1950

SPEC Heliconius clysonymus Latreille, [1817]

SUBS Heliconius clysonymus clysonymus Latreille, [1817]
SUBS Heliconius clysonymus hygiana (Hewitson, 1867)
SUBS Heliconius clysonymus montanus Salvin, 1871
SUBS Heliconius clysonymus tabaconas K. Brown, 1976

SPEC Heliconius congener Weymer, 1890

SUBS Heliconius congener congener Weymer, 1890
SUBS Heliconius congener aquilionaris K. Brown, 1976
SUBS Heliconius congener ocannensis Stichel, 1905

SPEC Heliconius cydno (E. Doubleday, 1847)

SUBS Heliconius cydno cydno (E. Doubleday, 1847)
SUBS Heliconius cydno alithea (Hewitson, 1869)
SUBS Heliconius cydno barinasensis Masters, 1973
SUBS Heliconius cydno chioneus Bates, 1864
SUBS Heliconius cydno cordula Neustetter, 1913
SUBS Heliconius cydno cydnides Staudinger, 1885
SUBS Heliconius cydno gadouae K. Brown & F. Fernández, 1985
SUBS Heliconius cydno galanthus Bates, 1864
SUBS Heliconius cydno hermogenes (Hewitson, [1858])
SUBS Heliconius cydno lisethae Neukirchen, 1995
SUBS Heliconius cydno wanningeri Neukirchen, 1991
SUBS Heliconius cydno weymeri Staudinger, 1897
SUBS Heliconius cydno zelinde Butler, 1869

SPEC Heliconius pachinus Salvin, 1871
SPEC Heliconius demeter Staudinger, 1897

SUBS Heliconius demeter demeter Staudinger, 1897
SUBS Heliconius demeter angeli Neukirchen, 1997
SUBS Heliconius demeter beebei J. Turner, 1966
SUBS Heliconius demeter bouqueti Nöldner, 1901
SUBS Heliconius demeter karinae Neukirchen, 1990
SUBS Heliconius demeter neildi Neukirchen, 1997
SUBS Heliconius demeter tambopata Lamas, 1985
SUBS Heliconius demeter terrasanta K. Brown & Benson, 1975
SUBS Heliconius demeter titan Neukirchen, 1995
SUBS Heliconius demeter turneri K. Brown & Benson, 1975
SUBS Heliconius demeter ulysses K. Brown & Benson, 1975
SUBS Heliconius demeter zikani K. Brown & Benson, 1975
SUBS Heliconius demeter joroni Lamas & Rosser, 2018

SPEC Heliconius eratosignis (Joicey & Talbot, 1925)

SUBS Heliconius eratosignis eratosignis (Joicey & Talbot, 1925)
SUBS Heliconius eratosignis ucayalensis H. Holzinger & R. Holzinger, 1975

SPEC Heliconius doris (Linnaeus, 1771)

SUBS Heliconius doris viridis Staudinger, 1885
SUBS Heliconius doris doris (Linnaeus, 1771)
SUBS Heliconius doris dives (Oberthür, 1920)
SUBS Heliconius doris obscurus Weymer, 1891

SPEC Heliconius egeria (Cramer, 1775)

SUBS Heliconius egeria egeria (Cramer, 1775)
SUBS Heliconius egeria christiani Neukirchen, 1997
SUBS Heliconius egeria egerides Staudinger, 1897
SUBS Heliconius egeria homogena Bryk, 1953
SUBS Heliconius egeria hyas Weymer, 1883
SUBS Heliconius egeria keithbrowni Neukirchen, 1992
SUBS Heliconius egeria mariasibyllae Neukirchen, 1991

SPEC Heliconius eleuchia (Hewitson, [1854])

SUBS Heliconius eleuchia eleuchia (Hewitson, [1854])
SUBS Heliconius eleuchia eleusinus Staudinger, 1885
SUBS Heliconius eleuchia primularis Butler, 1869

SPEC Heliconius elevatus Nöldner, 1901

SUBS Heliconius elevatus elevatus Nöldner, 1901
SUBS Heliconius elevatus bari (Oberthür, 1902)
SUBS Heliconius elevatus lapis Lamas, 1976
SUBS Heliconius elevatus perchlora Joicey & Kaye, 1917
SUBS Heliconius elevatus pseudocupidineus Neustetter, 1931
SUBS Heliconius elevatus roraima J. Turner, 1966
SUBS Heliconius elevatus schmassmanni Joicey & Talbot, 1925
SUBS Heliconius elevatus sonjae Neukirchen, 1997
SUBS Heliconius elevatus taracuanus Bryk, 1953
SUBS Heliconius elevatus tumatumari Kaye, 1906
SUBS Heliconius elevatus willmotti Neukirchen, 1997
SUBS Heliconius elevatus zoelleri Neukirchen, 1990

SPEC Heliconius erato (Linnaeus, 1758)

SUBS Heliconius erato erato (Linnaeus, 1758)
SUBS Heliconius erato adana J. Turner, 1967
SUBS Heliconius erato amalfreda Riffarth, 1901
SUBS Heliconius erato amazona Staudinger, 1897
SUBS Heliconius erato amphitrite Riffarth, 1901
SUBS Heliconius erato chestertonii (Hewitson, 1872)
SUBS Heliconius erato colombina Staudinger, 1897
SUBS Heliconius erato cruentus Lamas, 1998
SUBS Heliconius erato cyrbia (Godart, 1819)
SUBS Heliconius erato demophoon Ménétriés, 1855
SUBS Heliconius erato dignus Stichel, 1923
SUBS Heliconius erato emma Riffarth, 1901
SUBS Heliconius erato estrella Bates, 1862
SUBS Heliconius erato etylus Salvin, 1871
SUBS Heliconius erato favorinus Hopffer, 1874
SUBS Heliconius erato guarica Reakirt, 1868
SUBS Heliconius erato hydara (Hewitson, 1867)
SUBS Heliconius erato lativitta Butler, 1877
SUBS Heliconius erato lichyi K. Brown & F. Fernández, 1985
SUBS Heliconius erato luscombei Lamas, 1976
SUBS Heliconius erato magnifica Riffarth, 1900
SUBS Heliconius erato microclea Kaye, 1907
SUBS Heliconius erato notabilis Salvin & Godman, 1868
SUBS Heliconius erato petiverana (E. Doubleday, 1847)
SUBS Heliconius erato phyllis (Fabricius, 1775)
SUBS Heliconius erato reductimacula Bryk, 1953
SUBS Heliconius erato tobagoensis Barcant, 1982
SUBS Heliconius erato venus Staudinger, 1882
SUBS Heliconius erato venustus Salvin, 1871

SPEC Heliconius ethilla (Godart, 1819)

SUBS Heliconius ethilla ethilla (Godart, 1819)
SUBS Heliconius ethilla adela Neustetter, 1912
SUBS Heliconius ethilla aerotome C. Felder & R. Felder, 1862
SUBS Heliconius ethilla cephallenia C. Felder & R. Felder, 1865
SUBS Heliconius ethilla chapadensis K. Brown, 1973
SUBS Heliconius ethilla claudia Godman & Salvin, 1881
SUBS Heliconius ethilla eucoma (Hübner, [1831])
SUBS Heliconius ethilla flavofasciatus Weymer, 1894
SUBS Heliconius ethilla flavomaculatus Weymer, 1894
SUBS Heliconius ethilla hyalina Neustetter, 1928
SUBS Heliconius ethilla jaruensis K. Brown, 1976
SUBS Heliconius ethilla latona Neukirchen, 1998
SUBS Heliconius ethilla mentor Weymer, 1883
SUBS Heliconius ethilla metalilis Butler, 1873
SUBS Heliconius ethilla michaelianius Lamas, 1988
SUBS Heliconius ethilla narcaea (Godart, 1819)
SUBS Heliconius ethilla nebulosa Kaye, 1916
SUBS Heliconius ethilla neukircheni Lamas, 1998
SUBS Heliconius ethilla numismaticus Weymer, 1894
SUBS Heliconius ethilla penthesilea Neukirchen, 1994
SUBS Heliconius ethilla polychrous C. Felder & R. Felder, 1865
SUBS Heliconius ethilla semiflavidus Weymer, 1894
SUBS Heliconius ethilla thielei Riffarth, 1900
SUBS Heliconius ethilla tyndarus Weymer, 1897
SUBS Heliconius ethilla yuruani K. Brown & F. Fernández, 1985

SPEC Heliconius hecale (Fabricius, 1776)

SUBS Heliconius hecale hecale (Fabricius, 1776)
SUBS Heliconius hecale anderida (Hewitson, [1853])
SUBS Heliconius hecale annetta Riffarth, 1900
SUBS Heliconius hecale australis K. Brown, 1976
SUBS Heliconius hecale barcanti K. Brown, 1976
SUBS Heliconius hecale clearei A. Hall, 1930
SUBS Heliconius hecale discomaculatus Weymer, 1891
SUBS Heliconius hecale ennius Weymer, 1891
SUBS Heliconius hecale felix Weymer, 1894
SUBS Heliconius hecale fornarina (Hewitson, 1854)
SUBS Heliconius hecale holcophorus Staudinger, 1897
SUBS Heliconius hecale humboldti Neustetter, 1928
SUBS Heliconius hecale ithaca C. Felder & R. Felder, 1862
SUBS Heliconius hecale latus Riffarth, 1900
SUBS Heliconius hecale melicerta Bates, 1866
SUBS Heliconius hecale metellus Weymer, 1894
SUBS Heliconius hecale naxos Neukirchen, 1998
SUBS Heliconius hecale nigrofasciatus Weymer, 1894
SUBS Heliconius hecale novatus Bates, 1867
SUBS Heliconius hecale paraensis Riffarth, 1900
SUBS Heliconius hecale paulus Neukirchen, 1998
SUBS Heliconius hecale quitalena (Hewitson, [1853])
SUBS Heliconius hecale rosalesi K. Brown & F. Fernández, 1976
SUBS Heliconius hecale shanki Lamas & K. Brown, 1976
SUBS Heliconius hecale sisyphus Salvin, 1871
SUBS Heliconius hecale sulphureus Weymer, 1894
SUBS Heliconius hecale vetustus Butler, 1873
SUBS Heliconius hecale zeus Neukirchen, 1995
SUBS Heliconius hecale zuleika (Hewitson, 1854)

SPEC Heliconius hecalesia (Hewitson, [1854])

SUBS Heliconius hecalesia hecalesia (Hewitson, [1854])
SUBS Heliconius hecalesia eximius Stichel, 1923
SUBS Heliconius hecalesia formosus Bates, 1866
SUBS Heliconius hecalesia gynaesia (Hewitson, 1875)
SUBS Heliconius hecalesia longarena (Hewitson, 1875)
SUBS Heliconius hecalesia octavia Bates, 1866
SUBS Heliconius hecalesia romeroi K. Brown & F. Fernández, 1985

SPEC Heliconius hecuba (Hewitson, [1858])

SUBS Heliconius hecuba hecuba (Hewitson, [1858])
SUBS Heliconius hecuba bonplandi Neukirchen, 1991
SUBS Heliconius hecuba cassandra C. Felder & R. Felder, 1862
SUBS Heliconius hecuba choarina (Hewitson, 1872)
SUBS Heliconius hecuba creusa H. Holzinger & R. Holzinger, 1989
SUBS Heliconius hecuba crispus Staudinger, 1885
SUBS Heliconius hecuba flava K. Brown, 1979
SUBS Heliconius hecuba lamasi Neukirchen, 1991
SUBS Heliconius hecuba salazari Neukirchen, 1993
SUBS Heliconius hecuba tolima Fassl, 1912
SUBS Heliconius hecuba walteri Salazar, 1998

SPEC Heliconius hermathena (Hewitson, [1854])

SUBS Heliconius hermathena hermathena (Hewitson, [1854])
SUBS Heliconius hermathena duckei K. Brown & Benson, 1977
SUBS Heliconius hermathena renatae K. Brown & F. Fernández, 1985
SUBS Heliconius hermathena sabinae Neukirchen, 1992
SUBS Heliconius hermathena sheppardi K. Brown & Benson, 1977
SUBS Heliconius hermathena vereatta Stichel, 1912
SUBS Heliconius hermathena curua Freitas & Ramos, 2019

SPEC Heliconius heurippa (Hewitson, [1854])
SPEC Heliconius hierax (Hewitson, 1869)
SPEC Heliconius himera (Hewitson, 1867)
SPEC Heliconius hortense (Guérin-Méneville, [1844])
SPEC Heliconius ismenius Latreille, [1817]

SUBS Heliconius ismenius ismenius Latreille, [1817]
SUBS Heliconius ismenius boulleti Neustetter, 1928
SUBS Heliconius ismenius clarescens Butler, 1875
SUBS Heliconius ismenius fasciatus Godman & Salvin, 1877
SUBS Heliconius ismenius metaphorus Weymer, 1883
SUBS Heliconius ismenius occidentalis Neustetter, 1928
SUBS Heliconius ismenius telchinia (E. Doubleday, 1847)
SUBS Heliconius ismenius tilletti K. Brown & F. Fernández, 1976

SPEC Heliconius lalitae Brévignon, 1996
SPEC Heliconius leucadia Bates, 1862

SUBS Heliconius leucadia leucadia Bates, 1862
SUBS Heliconius leucadia andromeda Neukirchen, 1996
SUBS Heliconius leucadia birgitae Neukirchen, 1996
SUBS Heliconius leucadia pseudorhea Staudinger, 1897

SPEC Heliconius luciana Lichy, 1960

SUBS Heliconius luciana luciana Lichy, 1970
SUBS Heliconius luciana watunna Lichy, 1960

SPEC Heliconius melpomene (Linnaeus, 1758)

SUBS Heliconius melpomene melpomene (Linnaeus, 1758)
SUBS Heliconius melpomene aglaope C. Felder & R. Felder, 1862
SUBS Heliconius melpomene amandus Grose-Smith & W. F. Kirby, 1892
SUBS Heliconius melpomene amaryllis C. Felder & R. Felder, 1862
SUBS Heliconius melpomene anduzei K. Brown & F. Fernández, 1985
SUBS Heliconius melpomene bellula J. Turner, 1971
SUBS Heliconius melpomene burchelli Poulton, 1910
SUBS Heliconius melpomene cythera (Hewitson, 1869)
SUBS Heliconius melpomene ecuadorensis Emsley, 1964
SUBS Heliconius melpomene euryades Riffarth, 1900
SUBS Heliconius melpomene flagrans Stichel, 1919
SUBS Heliconius melpomene intersectus Neustetter, 1928
SUBS Heliconius melpomene madeira N. Riley, 1919
SUBS Heliconius melpomene malleti Lamas, 1988
SUBS Heliconius melpomene martinae Cast & Le Crom, 2012
SUBS Heliconius melpomene meriana J. Turner, 1967
SUBS Heliconius melpomene michellae Neukirchen, 1997
SUBS Heliconius melpomene nanna Stichel, 1899
SUBS Heliconius melpomene penelope Staudinger, 1894
SUBS Heliconius melpomene plesseni Riffarth, 1907
SUBS Heliconius melpomene pyrforus Kaye, 1907
SUBS Heliconius melpomene rosina (Boisduval, 1870)
SUBS Heliconius melpomene schunkei Lamas, 1976
SUBS Heliconius melpomene tessa Barcant, 1982
SUBS Heliconius melpomene thelxiope (Hübner, [1806])
SUBS Heliconius melpomene thelxiopeia Staudinger, 1897
SUBS Heliconius melpomene vicina Ménétriés, 1857
SUBS Heliconius melpomene vulcanus Butler, 1865
SUBS Heliconius melpomene xenoclea (Hewitson, [1853])

SPEC Heliconius nattereri C. Felder & R. Felder, 1865
SPEC Heliconius numata (Cramer, 1780)

SUBS Heliconius numata numata (Cramer, 1780)
SUBS Heliconius numata arcuella H. Druce, 1874
SUBS Heliconius numata aristiona (Hewitson, [1853])
SUBS Heliconius numata aulicus Weymer, 1883
SUBS Heliconius numata aurora Bates, 1862
SUBS Heliconius numata bicoloratus Butler, 1873
SUBS Heliconius numata ethra (Hübner, [1831])
SUBS Heliconius numata euphone C. Felder & R. Felder, 1862
SUBS Heliconius numata geminatus Weymer, 1894
SUBS Heliconius numata holzingeri F. Fernández & K. Brown, 1976
SUBS Heliconius numata ignotus Joicey & Kaye, 1917
SUBS Heliconius numata illustris Weymer, 1894
SUBS Heliconius numata jiparanaensis Neustetter, 1931
SUBS Heliconius numata lenaeus Weymer, 1891
SUBS Heliconius numata lyrcaeus Weymer, 1891
SUBS Heliconius numata mavors Weymer, 1894
SUBS Heliconius numata messene C. Felder & R. Felder, 1862
SUBS Heliconius numata mirus Weymer, 1894
SUBS Heliconius numata nubifer Butler, 1875
SUBS Heliconius numata peeblesi Joicey & Talbot, 1925
SUBS Heliconius numata pratti Joicey & Kaye, 1917
SUBS Heliconius numata robigus Weymer, 1875
SUBS Heliconius numata silvana (Stoll, 1781)
SUBS Heliconius numata sourensis K. Brown, 1976
SUBS Heliconius numata superioris Butler, 1875
SUBS Heliconius numata talboti Joicey & Kaye, 1917
SUBS Heliconius numata tarapotensis Riffarth, 1901
SUBS Heliconius numata zobrysi Fruhstorfer, 1910

SPEC Heliconius pardalinus Bates, 1862

SUBS Heliconius pardalinus pardalinus Bates, 1862
SUBS Heliconius pardalinus ariadne Neukirchen, 1995
SUBS Heliconius pardalinus butleri K. Brown, 1976
SUBS Heliconius pardalinus dilatus Weymer, 1894
SUBS Heliconius pardalinus julia Neukirchen, 2000
SUBS Heliconius pardalinus lucescens Weymer, 1894
SUBS Heliconius pardalinus maeon Weymer, 1891
SUBS Heliconius pardalinus orteguaza K. Brown, 1976
SUBS Heliconius pardalinus radiosus Butler, 1873
SUBS Heliconius pardalinus sergestus Weymer, 1894
SUBS Heliconius pardalinus tithoreides Staudinger, 1900

SPEC Heliconius peruvianus (C. Felder & R. Felder, 1859)
SPEC Heliconius ricini (Linnaeus, 1758)

SUBS Heliconius ricini ricini (Linnaeus, 1758)
SUBS Heliconius ricini insulanus (Stichel, 1909)

SPEC Heliconius sapho (Drury, 1782)

SUBS Heliconius sapho sapho (Drury, 1782)
SUBS Heliconius sapho candidus K. Brown, 1976
SUBS Heliconius sapho chocoensis K. Brown & Benson, 1975
SUBS Heliconius sapho leuce (E. Doubleday, 1847)

SPEC Heliconius hewitsoni (Hewitson, 1875)
SPEC Heliconius sara (Fabricius, 1793)

SUBS Heliconius sara sara (Fabricius, 1793)
SUBS Heliconius sara apseudes (Hübner, [1813])
SUBS Heliconius sara brevimaculata Riffarth, 1901
SUBS Heliconius sara elektra Neukirchen, 1998
SUBS Heliconius sara fulgidus Stichel, 1906
SUBS Heliconius sara magdalena Bates, 1864
SUBS Heliconius sara sprucei Bates, 1864
SUBS Heliconius sara theudela (Hewitson, 1874)
SUBS Heliconius sara veraepacis Bates, 1864
SUBS Heliconius sara williami Neukirchen, 1994

SPEC Heliconius telesiphe (E. Doubleday, 1847)

SUBS Heliconius telesiphe telesiphe (E. Doubleday, 1847)
SUBS Heliconius telesiphe cretacea Neustetter, 1916
SUBS Heliconius telesiphe sotericus Salvin, 1871

SPEC Heliconius timareta (Hewitson, 1867)

SUBS Heliconius timareta timareta (Hewitson, 1867)
SUBS Heliconius timareta timoratus Lamas, 1998

SPEC Heliconius tristero A. V.Z. Brower, 1996
SPEC Heliconius wallacei Reakirt, 1866

SUBS Heliconius wallacei wallacei Reakirt, 1866
SUBS Heliconius wallacei araguaia K. Brown, 1976
SUBS Heliconius wallacei colon Weymer, 1891
SUBS Heliconius wallacei flavescens Weymer, 1891
SUBS Heliconius wallacei kayei Neustetter, 1929
SUBS Heliconius wallacei mimulinus Butler, 1873

SPEC Heliconius xanthocles Bates, 1862

SUBS Heliconius xanthocles xanthocles Bates, 1862
SUBS Heliconius xanthocles buechei Neukirchen, 1992
SUBS Heliconius xanthocles cleoxanthe H. Holzinger & R. Holzinger, 1972
SUBS Heliconius xanthocles donatia Fruhstorfer, 1910
SUBS Heliconius xanthocles explicata K. Brown, 1976
SUBS Heliconius xanthocles hippocrene H. Holzinger & K. Brown, 1982
SUBS Heliconius xanthocles melete C. Felder & R. Felder, 1865
SUBS Heliconius xanthocles melior Staudinger, 1897
SUBS Heliconius xanthocles melittus Staudinger, 1897
SUBS Heliconius xanthocles napoensis H. Holzinger & K. Brown, 1982
SUBS Heliconius xanthocles paraplesius Bates, 1867
SUBS Heliconius xanthocles quindecim Lamas, 1976
SUBS Heliconius xanthocles rindgei H. Holzinger & K. Brown, 1982
SUBS Heliconius xanthocles similatus J. Zikán, 1937
SUBS Heliconius xanthocles vala Staudinger, 1885
SUBS Heliconius xanthocles zamora H. Holzinger & K. Brown, 1982


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